What kind of print defect is this?

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This “uninked” line is present on both sides of the note and is visible under backlight:

 

   

 

What caused this?

Wanted: Cambodia 2000 Riels 2007 P#59b (printed 2015) UNC or AU
https://www.linkedin.com/in/brandon-bertolli-b6500522/recent-activity/all/

A small fold in the paper?

Just call me Bram

No new swaps for the moment, still too many half-ongoing swaps to clean up!

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